The Athlone man continues as Andy Farrell's first choice outside centre.
Robbie Henshaw will partner Bundee Aki at centre in the Ireland team against South Africa in Saturday’s first Test at Loftus Versfeld Stadium.
Jamie Osborne makes his international debut at full back in the absence of Hugo Keenan who has committed his summer to Ireland Olympic Sevens effort.
While Craig Casey will partner his provincial teammate Jack Crowley at half back.
Peter O'Mahoney will once again captain the side from blindside flanker with Josh Van Der Flier and Caelon Doris joining him in the back row.
Our line up to face South Africa in the first Test! 🟢#TeamOfUs
— Irish Rugby (@IrishRugby) July 4, 2024
Saturday's contest in Stellenbosch kicks off at 4pm.